et al., 2000). The p
Silencer
expression plasmids included GFP for identification of
transfected cells. Silencing of SytV and Syt IX levels did not
change basal secretion. However, glucose-induced insulin
release was decreased suggesting that inhibition may be by
impairment of the exocytotic process and that Syt V and Syt IX
act as positive modulators of exocytosis. The addition of the
phorbol ester PMA, which potentiates insulin exocytosis distal
to the increase of cytosolic Ca
2+ showed that both
proteins are directly involved in the
Ca
2+-dependent stimulation of exocytosis. The
association of Syt IX with insulin granules in β-cells
suggests that Syt IX may be the Ca
2+ sensor for
insulin secretion in β-cells.
References
Chapman ER (2002) Synaptotagmin a Ca 2+
sensor that triggers exocytosis? Nat Rev Mol Cell Biol
3:498-508.
Gut A, Kiraly CE, Fukuda M, Mikoshiba K,
Wollheim CB, Lang J (2001) Expression and localisation of
synaptotagmin isoforms in endocrine beta-cells: their function
in insulin exocytosis. J Cell Sci 114:1709-1716.
Iezzi M, Regazzi R, Wollheim, CB (2000)
The Rab3-interacting molecule RIM is expressed in pancreatic
beta-cells and is implicated in insulin exocytosis. FEBS
Lett 26:66-70.
Nagamatsu S, Nakamichi Y, Yamamura C,
Matsushima S, Watanabe T, Ozawa S, Furukawa H, Ishida H
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